Karl Andrew Johnson


Department of Biology                                                                   kjohnson@haverford.edu

Haverford College                                                                                      (T) 610-896-1306

370 Lancaster Ave.                                                                                    (F) 610-896-4963

Haverford PA 19041                                                  haverford.edu/biology/johnson



Professional Experience


2007-present    Professor, Department of Biology, Haverford College

2009-2012       Chair, Department of Biology, Haverford College

2001-2004       Chair, Department of Biology, Haverford College

1999-2007       Associate Professor, Department of Biology, Haverford College

1993-1999       Assistant Professor, Department of Biology, Haverford College

1992                Visiting Assistant Professor, Yale College (Summer Programs)

1992                Visiting Assistant Professor, Connecticut College (Spring Semester)

1991                Lecturer in Biology, Yale University (Fall Semester)




1989-1993       Postdoctoral Fellow, Yale University

                                    Advisor: Joel Rosenbaum, Ph.D.

1989                Ph.D. in Biology, Yale University

Advisor: Ian Sussex, Ph.D.

1987                M.Phil. in Biology, Yale University

                                    Advisor: Mary Helen Goldsmith. Ph.D.

1983                B.A. in Biology, Swarthmore College

                                    Senior Research Advisor: Mark Jacobs, Ph.D.


Grants (1999-present)


2000-2005       National Science Foundation RUI Research Grant

                                    Phage-Based Components for Nanoscale Assembly

$180,000 (to K.A.J.)

2000-2005       National Science Foundation MRI Instrumentation Grant

                                    Acquisition of Molecular and Cellular Imaging Instrumentation

$996,294 (co-PI)

2000-2005       National Science Foundation RUI Research Grant

                                    Tubulin Folding and Assembly in Chlamydomonas

$247,000 (MCB-9982733 to K.A.J.)

2000-2006       David and Lucille Packard Foundation Grant

                                    Protein-Based Biomaterials for Nanotechnology

$966,020 (co-PI)

2000-2002       National Science Foundation Instrumentation Grant

RUI: Advanced Microscopy and Manipulation Cluster for Biological and Biophysical Studies.  $169,271 (DBI-0070361, co-PI)

2000-2002       National Science Foundation Instrumentation Grant

A Confocal Microscope for Research in the Biological Sciences

$165,985 (DBI-0070046, co-PI)

1999-2000       Zimmer Corporation Instrumentation Grant

                                    Atomic Force Microscopy $100,000 (co-PI)

1995-1999       National Science Foundation RUI Research Grant

Characterization of a Molecular Chaperone in the Eukaryotic Flagellum $240,000 (MCB-9506236 to K.A.J.)

And participation in program grants to Haverford College from the Howard Hughes Medical Institute, the Merck Foundation, the Sherman Fairchild Foundation, and the Beckman Foundation

Awards, Honors and Fellowships (1999-present)


2004-2005       Accrued Sabbatical leave (half-time)

2001-2002       Teaching with Technology Development Award, Haverford College

2000-2001       Post-Tenure Leave (full time)

1999                Faculty Innovation in Teaching Award, Haverford College


Courses Taught at Haverford (1999-present)


HBiol 123        Perspectives in Biology: Scientific Literacy- a class for nonscientists

HBiol 200        Cell Structure and Function – a year long, team-taught gateway to the major

HBiol 300        Laboratory in Biochemistry/Molecular Biology – intensive co-taught junior laboratory

HBiol 302        Cell Architecture - a junior-level cell biology lecture course

HBiol 351        Molecular Motors and Biological Nanomachines - a literature-based senior seminar

HBiol 407        Senior Research Tutorial – senior research theses (required for the major)

HBiol 499        Senior Departmental Studies – a senior collective experience


Publications from Haverford (* denotes undergraduate co-authors)


Kurtuldu H, Guasto JS, Johnson KA, Gollub JP. (2011) Enhancement of biomixing by swimming algal cells in two-dimensional films. Proc Natl Acad Sci U S A. 2011 Jun 28;108(26):10391-5. Epub 2011 Jun 9. PMID: 21659630

Guasto JS, Johnson KA, Gollub JP. (2010) Oscillatory flows induced by microorganisms swimming in two dimensions. Phys Rev Lett. 105(16):168102. PMID: 21231018

Kokona B, *Kim AM, *Roden RC, *Daniels JP, *Pepe-Mooney BJ, *Kovaric BC, de Paula JC, Johnson KA, Fairman R. (2009) Self Assembly of Coiled-Coil Peptide-Porphyrin Complexes. Biomacromolecules. 2009 Apr 17. PMID: 19374349

*Shapiro J, *Ingram J, Johnson KA (2005) Characterization of a molecular chaperone present in the eukaryotic flagellum.  Eukaryot Cell. 4(9):1591-4.  PMID: 16151252

*Rigotti DJ, Kokona B, *Horne T, *Acton EK, *Lederman CD, Johnson KA, Manning RS, Kane SA, Smith WF, Fairman R.  (2005) Quantitative atomic force microscopy image analysis of unusual filaments formed by the Acanthamoeba castellanii myosin II rod domain. Anal Biochem. 346(2):189-200. PMID: 16213459

Fossella J, Samant SA, Silver LM, King SM, Vaughan KT, Olds-Clarke P, Johnson KA, Mikami A, Vallee RB, Pilder SH. (2000) An axonemal dynein at the Hybrid Sterility 6 locus: implications for t haplotype-specific male sterility and the evolution of species barriers.  Mamm Genome. 11(1):8-15.  PMID: 10602986

Johnson KA. (1998) The axonemal microtubules of the Chlamydomonas flagellum differ in tubulin isoform content.  J Cell Sci. 1998 111( Pt 3):313-20.  PMID: 9427680

*Bloch MA, Johnson KA. (1995) Identification of a molecular chaperone in the eukaryotic flagellum and its localization to the site of microtubule assembly. J Cell Sci. 108 ( Pt 11):3541-5. PMID: 8586665

Johnson, K.A. (1995) Keeping the beat: form meets function in the Chlamydomonas flagellum.  BioEssays 17:847-854.  DOI: 10.1002/bies.950171006

Johnson KA. (1995) Immunoelectron microscopy. Methods Cell Biol. 1995;47:153-62. PMID: 7476481

Bernstein M, Beech PL, Johnson KA, Kozminski KG and Rosenbaum JL. (1995) New motilities and motors in the flagella of Chlamydomonas. In: 45th Colloquium Mosbach: The Cytoskeleton, ed. by B. Jockusch, E. Mandelkow and K. Weber. Springer-Verlag, Heidelberg, Germany, pp. 11-21.

Johnson KA, Haas MA, Rosenbaum JL. (1994) Localization of a kinesin-related protein to the central pair apparatus of the Chlamydomonas reinhardtii flagellum.  J Cell Sci. 107 ( Pt 6):1551-6. PMID: 7962197


Publications prior to Haverford


Kozminski KG, Johnson KA, Forscher P, Rosenbaum JL. (1993) A motility in the eukaryotic flagellum unrelated to flagellar beating. Proc Natl Acad Sci U S A. 90(12):5519-23. PMID: 8516294

Johnson KA, Rosenbaum JL. (1993) Flagellar regeneration in Chlamydomonas: a model system for studying organelle assembly.  Trends Cell Biol. 3(5):156-61.  PMID: 14731610

Johnson KA, Rosenbaum JL. (1992) Polarity of flagellar assembly in Chlamydomonas. J Cell Biol. 119(6):1605-11. PMID: 1281816

Johnson KA, Rosenbaum JL. (1992) Replication of basal bodies and centrioles. Curr Opin Cell Biol. 4(1):80-5. Review. PMID: 1558756

Johnson KA, Rosenbaum JL. (1991) Basal bodies and DNA. Trends Cell Biol. 1(6):145-9. PMID: 14731856

Diener DR, Curry AM, Johnson KA, Williams BD, Lefebvre PA, Kindle KL, Rosenbaum JL. Rescue of a paralyzed-flagella mutant of Chlamydomonas by transformation. Proc Natl Acad Sci U S A. 87(15):5739-43. PMID: 2377611

Johnson KA, Rosenbaum JL. (1990) Basal bodies of Chlamydomonas reinhardtii do not contain immunologically detectable DNA. Cell. 62(4):615-9. PMID: 2386994

Johnson KA, Sussex IM. (1990) Genomic amplification in the cotyledon parenchyma of common bean. Chromosoma. 99(3):223-30. PMID: 2397660

Johnson, KA. (1989) The cytologic and molecular analysis of the developmentally controlled increase in nuclear DNA content in the cotyledon parenchyma of common bean, Phaseolus vulgaris.  Ph.D. Dissertation, Yale University, New Haven, CT.


Synergistic Activities (1999-present)


Ad Hoc Search Committees

                        Instrument Specialies Search, 2011

Economics Search Chair, 2006-2007

Biochemistry Search (at Bryn Mawr College) 2006-2007

                        Physical Chemistry Search Chair, 2005-2006

Molecular and Cellular Evolution Search, 2003-2004

                        Ecology Search (at Bryn Mawr College) 2002-2003

Biology/Chemistry Business Manager Search, 2002

Molecular Neurobiology Search, 2001-2002

                        Science Librarian Search, 2000

                        Biology Search, 1998-1999

Histology/microscopy workshop, Garnet Valley High School AP Biology Class, 2005

Committee for Environmental Responsibility, Haverford College, 2005-2008

First Interdisciplinary Science Summer Journal Club, Haverford College, organizer, 2003

Howard Hughes Program Coordinating Committee 2002-2004

Institutional Review Board (Human Subjects), Haverford College, 2002-2004

Koshland Integrated Natural Sciences Center Steering Committee, 2002-2003

Radiation Safety Officer, Haverford College, 1999-2000

Beckman Scholars Selection Committee, 1999-2000

Distinguished Visitors Committee, 1997-2000 (Chair)


Faculty Advisor to First and Second Year Students, and Majors (annually)

Biology Department Internal Committees (Equipment, Building, Web Admin., etc.)

Ad Hoc Reviewer (Grant Applications): National Science Foundation, National Institutes of Health (Minority Biomedical Research Support Program)

Ad Hoc Reviewer (Manuscripts): Genomics, Trends in Cell Biology, Journal of Cell Science, Chromosoma

Ad Hoc Reviewer/Glossator, Personnel Cases: Haverford, Bryn Mawr, Swarthmore, Reed and Loyola Colleges

Invited contributor, Graduate Record Examination in Biochemistry, Cell and Molecular Biology


Professional Memberships


American Association for the Advancement of Science

American Society for Cell Biology

American Society for Microbiology                                                                


updated 070629 KJ